Initialize_AnyThread and CacheBones_AnyThread can be called repeatedly on sub-graphs below Saved Pose nodes when they are part of a linked anim graph and the main anim graph has yet to be initialized. This can happen when bTickAnimationOnSkeletalMeshInit is disabled, and LinkAnimClassLayers is called on the first frame that a mesh is active. LinkAnimClassLayers results in a call to Initialize_AnyThread and CacheBones_AnyThread on nodes within the linked graph's anim graph. But the synchronization counters are invalid as we use the counters from the parent graph (which hasn't been initialized yet).