Niagara Particles in Mesh Shape Don't Render in Scene or Editor Preview

Niagara particles aren't rendered in the Niagara Editor Preview or the scene when an emitter is in it.

Found in 4.24.1 CL#10756747
Reproduced in 4.25 Main CL#10808197

1. Enable the Niagara plugin.
2. Create a Niagara Emitter.
3. Remove all the modules in the Particle Update section.
4. Remove the "Spawn Rate" module in the Emitter update section.
5. Remove the "Calculate Size by Mass" & "Add Velocity" modules in the Particle Spawn section.
6. Add the "Sample Skeletal Mesh Surface" & "Skeletal Mesh Surface Location" modules in the Particle Spawn section.
7. Assign the SK_Mannequin mesh in the Default Mesh field in "Sample Skeletal MeshSurface".
8. Create a niagara system from the emitter.
9. Place the niagara system in the scene.

Result: The niagara particles aren't rendered in both the Editor Preview and the scene.

Expected: The niagara particles render in the Editor Preview and the scene.